Note: Pdf files may not be downloaded for commercial purposes, only single copies for personal use are allowed.


Prins, H.H.T. and F. van Langevelde (eds.) (2008) Resource Ecology. Spatial and temporal aspects of foraging. Springer, Dordrecht, The Netherlands, 304 p. (link)

Peer reviewed


153. Van Dooremalen, C. and Frank van Langevelde (2021) Can colony size of honeybees (Apis mellifera) be used as predictor for colony losses due to Varroa destructor during winter? Agriculture 11:529

152. Kauffman, M.J., F. Cagnacci, S. Chamaillé-Jammes, M. Hebblewhite, J.G.C. Hopcraft, J.A. Merkle, T. Mueller, A. Mysterud, W. Peters, C. Roettger, A. Steingisser, J.E. Meacham, K. Abera, J. Adamczewski, E.O. Aikens, H. Bartlam-Brooks, E. Bennitt, J. Berger, C. Boyd, S.D. Côté, L. Debeffe, A.S. Dekrout, N. Dejid, E. Donadio, L. Dziba, W.F. Fagan, C. Fischer, S. Focardi, J.M. Fryxell, R.W.S. Fynn, C. Geremia, B.A. González, A. Gunn, E. Gurarie, M. Heurich, J. Hilty, M. Hurley, A. Johnson, K. Joly, P. Kaczensky, C.J. Kendall, P. Kochkarev, L. Kolpaschikov, R. Kowalczyk, F. van Langevelde, B.V. Li, A.L. Lobora, A. Loison, T.H. Madiri, D. Mallon, P. Marchand, R.A. Medellin, E. Meisingset, E. Merrill, A.D. Middleton, K.L. Monteith, M. Morjan, T.A. Morrison, S. Mumme, R. Naidoo, A. Novaro, J.O. Ogutu, K.A. Olson, A. Oteng-Yeboah, R.J.A. Ovejero, N. Owen-Smith, A. Paasivaara, C. Packer, D. Panchenko, L. Pedrotti, A.J. Plumptre, C.M. Rolandsen, S. Said, A. Salemgareyev, A. Savchenko, P. Savchenko, H. Sawyer, M. Selebatso, M. Skroch, E. Solberg, J.A. Stabach, O. Strand, M.J. Suitor, Y. Tachiki, A. Trainor, A. Tshipa, M.Z. Virani, C. Vynne, S. Ward, G. Wittemyer and W. Xu and S. Zuther (2021) Mapping out a future for ungulate migrations. Science 372:566–569

151. Chepkwony, R., S. van Bommel and F. van Langevelde (2021) Interactive effects of biological, human and environmental factors on tick loads in Boran cattle in tropical drylands. Parasites & Vectors 14:1–8

150. Veenendaal, E.M., M. Torello-Raventos, H.S. Miranda, N.M. Sato, T.A.J. Janssen, F. van Langevelde and J. Lloyd (2021) Fire regimes, fire experiments and alternative stable states in mesic savannas. A response to Laris & Jacobs (2021). New Phytologist 231:14–18

149. Issifu, H., P. Vergeer, G.K.D. Ametsitsi, J. Klijn, P. Sartorelli, M. Tanson, H. Bayor, V. Logah, F. van Langevelde and E.M. Veenendaal (accepted) Responses of tropical tree seedlings in the forest-savanna boundary to combined effects of grass competition and fire. Biotropica

148. De Knegt, H.J., J.A.J. Eikelboom, F. van Langevelde, W.F. Spruyt and H.H.T. Prins (2021) Timely poacher detection and localization using sentinel animal movement. Scientific Reports 11:4596

147. Vissia, S., R. Wadhwa and F. van Langevelde (2021) Co-occurrence of high densities of brown hyena and spotted hyena in central Tuli, Botswana. Journal of Zoology 314:143–150

146. Okita-Ouma, B., F. van Langevelde, I.M.A. Heitkönig, P. Maina, S.E. van Wieren and Herbert H.T. Prins (2021) Relationships of reproductive performance indicators in black rhinoceros (Diceros bicornis michaeli) with plant available moisture, plant available nutrients and woody cover. African Journal of Ecology 59:2–16


145. Van Langevelde, F., F. Kiggen, C. van Dooremalen and B. Cornelissen (2020) Corpse removal increases when honey bee colonies experience high Varroa destructor infestation. Insectes Sociaux 67:507–513

144. Eikelboom, J.A.J., H.J. de Knegt, M. Klaver, F. van Langevelde, T. van der Wal and H.H.T. Prins (2020) Inferring an animal’s environment through biologging: quantifying the environmental contribution to animal movement. Movement Ecology 8:1–18

143. Chepkwony, R., C. Castagna, I.M.A. Heitkönig, S. van Bommel and F. van Langevelde (2020) Associations between monthly rainfall and mortality in cattle due to East Coast fever, anaplasmosis and babesiosis. Parasitology 147:1743–1751

142. Muijres, F.T., C. van Dooremalen, M. Lankheet, H. Lugt, L.J. de Vries and F. van Langevelde (2020) Varroa destructor infestation impairs the improvement of landing performance in foraging honeybees. Royal Society Open Science 7:201222

141. Ametsitsi, G.K.D., F. van Langevelde, V. Logah, T. Janssen, J.A. Medina-Vega, H. Issifu, L. Ollivier, K. den Hartogh, T. Adjei-Gyapong, S. Adu-Bredu, J. Lloyd and E.M. Veenendaal (2020) Fixed or mixed? Variation in tree functional types and vegetation structure in a forest-savanna ecotone in West Africa. Journal of Tropical Ecology 36:133–149

140. Eikelboom, J.A.J., R.J.M. Nuijten, Y.X.G. Wang, B. Schroder, F. van Langevelde and H.H.T. Prins (2020) Will legal international rhino horn trade save wild rhino populations? Global Ecology and Conservation 23:e01145

139. De Vries, L.J., F. van Langevelde, C. van Dooremalen, I.G. Kornegoor, M.J. Lankheet, J.L. van Leeuwen, M. Naguib and F.T. Muijres (2020) Bumblebees land remarkably well in red-blue greenhouse LED light conditions. Biology Open 

138. Owen-Smith, N., G. Hopcraft, T. Morrison, S. Chamaillé-Jammes, R. Hetem, E. Bennitt and F. van Langevelde (2020) Movement ecology of large herbivores in African savannas: current knowledge and gaps. Mammal Review 50:252–266

137. Bastiaansen, R., A. Doelman, F. van Langevelde and V. Rottschäfer (2020) Modelling honey bee colonies in winter using a Keller-Segel model with a sign-changing chemotactic coefficient. SIAM Journal on Applied Mathematics 80:839–863

136. Boonman, C.C.F., F. van Langevelde, I. Oliveras, J. Couédon, N. Luijken, D. Martini and E.M. Veenendaal (2020) On the importance of root traits in seedlings of tropical tree species. New Phytologist 227:156167

135. Spikmans, F., P. Lemmers, H.J.M. op den Camp, E. van Haren, F. Kappen, A. Blaakmeer, G. van der Velde, F. van Langevelde, R.S.E.W. Leuven and T. van Alen (2020) Impact of the invasive alien topmouth gudgeon (Pseudorasbora parva) and its associated parasite Sphaerothecum destruens on native fish species. Biological Invasions 22:587–601

134. Van Langevelde, F., V. Comor, S. de Bie, H.H.T. Prins and M.P. Thakur (2020) Disturbance regulates the density – body mass relationship of soil fauna. Ecological Applications 30:e02019


133. Eikelboom, J.A.J., J. Wind, E. van de Ven, L.M. Kenana, B. Schroder, H.J. de Knegt, F. van Langevelde and H.H.T. Prins (2019) Improving the precision and accuracy of animal population estimates with aerial image object detection. Methods in Ecology and Evolution 10:1875–1887

132. Huang, Z.Y.X., C. Xu, F. van Langevelde, T. Langendoen, T. Mundkur, Y. Si, H. Tian, R.H.S. Kraus, M. Gilbert, G.-Z. Han, X. Ji, H.H.T. Prins and W.F. de Boer (2019) Contrasting effects of host species and phylogenetic diversity on the occurrence of HPAI H5N1 in European wild birds. Journal of Animal Ecology 88:1044–1053

131. Van Langevelde, F., C.A.D.M. van de Vijver, H.H.T. Prins and T.A. Groen (2019) Effects of grazing and browsing on tropical savanna vegetation. In: I.J. Gordon and H.HT Prins (eds.), The ecology of browsing and grazing II. Springer, Berlin, Germany, pp. 237–257

130. Issifu, H., G. Ametsitsi, L.J. de Vries, G.D. Djagbletey, S. Adu-Bredu, P. Vergeer, F. van Langevelde and E.M. Veenendaal (2019) Variation in vegetation cover and seedling performance of tree species in a forest-savanna ecotone. Journal of Tropical Ecology 35:74–82

129. Weterings, M.J.A., S.P. Ewert, J.N. Peereboom, H.J. Kuipers, D.P.J. Kuijper, H.H.T. Prins, P.A. Jansen, F. van Langevelde and S.E. van Wieren (2019) Implications of shared predation for space use in two sympatric leporids. Ecology and Evolution 9:3457–3469

128. Tomlinson, K.W., F.J. Sterck, E.R.M. Barbosa, S. de Bie, H.H.T. Prins and F. van Langevelde (2019) Seedling growth of savanna tree species from three continents under grass competition and nutrient limitation in a greenhouse experiment. Journal of Ecology 107:1051–1066


127. Armani, M., F. van Langevelde, K.W. Tomlinson, S. Adu-Bredu, G. Djagbletey and E.M. Veenendaal (2018) Compositional patterns of overstorey and understorey woody communities in a forest-savanna boundary in Ghana. Plant Ecology & Diversity 11:451-463

126. Wato, Y.A., H.H.T. Prins, I.M.A. Heitkonig, G. Wahungu, S. Ngene, S. Njumbi, F. van Langevelde (2018) Movement patterns of African elephants (Loxodonta africana) in a semi-arid savanna suggest that they have information on the location of dispersed water sources. Frontiers in Ecology and Evolution 6

125. Chepkwony, R., S. van Bommel and F. van Langevelde (2018) Citizen science for development: Potential role of mobile phones in information sharing on ticks and tick-borne diseases in Laikipia, Kenya. NJAS – Wageningen Journal of Life Sciences 86–87:123–135

124. Weterings, M.J.A., S. Moonen, H.H.T. Prins, S.E. van Wieren and F. van Langevelde (2018) Food quality and quantity is more important in explaining foraging of an intermediate-sized mammalian herbivore than predation risk or competition. Ecology and Evolution 8:8419–8432

123. Donners, M., R.H.A. van Grunsven, D. Groenendijk, F. van Langevelde, J.W. Bikker, T. Longcore and E.M. Veenendaal (2018) Colours of attraction: Modelling insect flight to light behaviour. Journal of Experimental Zoology Part A 329:434–440

122. Lameris, T., E. Kleyheeg, P.A. Jansen, J.S. Brown and F. van Langevelde (2018) Nest defensibility decreases home-range size in central-place foragers. Behavioural Ecology 29:1038–1045

121. De Vries, L.J. and F. van Langevelde (2018) Two different strategies of host manipulation allow parasites to persist in intermediate-definitive host systems. Journal of Evolutionary Biology 31:393–404

120. Van Langevelde, F., M. Braamburg-Annegarn, M.E. Huigens, R. Groendijk, O. Poitevin, J.R. van Deijk, W.N. Ellis, R.H.A. van Grunsven, R. de Vos, R.A. Vos, M. Franzén and M.F. WallisDeVries (2018) Declines in moth populations stress the need for conserving dark nights. Global Change Biology 24:925–932

119. De Schaetzen, F., F. van Langevelde and M.F. WallisDeVries (2018) The influence of wild boar (Sus scrofa) on microhabitat quality for the endangered butterfly Pyrgus malvae in the Netherlands. Journal of Insect Conservation 22:51–59

118. Veenendaal, E.M., M. Torello-Raventos, H.S. Miranda, N.M. Sato, I. Oliveras, F. van Langevelde, G.P. Asner and J. Lloyd (2018) On the relationship between fire regime and vegetation structure in the tropics. New Phytologist 218:153–166

117. Nkoba, K., F. van Langevelde and S.K. Raina (2018) Forest degradation influences nesting site selection of Afro-tropical stingless bee species in a tropical rain forest, Kenya. African Journal of Ecology 56:669–674


116. Groen, T.A., C.A.D.M. van de Vijver and F. van Langevelde (2017) Do spatially homogenising and heterogenising processes affect transitions between alternative stable states? Ecological Modelling 365:119–128

115. Panziera, D., F. van Langevelde and T. Blacquière (2017) Varroa Sensitive Hygiene contributes to naturally selected varroa resistance in honey bees. Journal of Apicultural Research 56:635642

114. Wynhoff, I. and F. van Langevelde (2017) Phengaris (Maculinea) teleius butterflies select host plants close to Myrmica ants for oviposition, but P. nausithous do not. Entomologia Experimentalis et Applicata 165:918

113. Kruitwagen, A., F. van Langevelde, C. van Dooremalen and T. Blacquière (2017) Naturally selected honey bee (Apis mellifera) colonies resistant to Varroa destructor do not groom more intensively. Journal of Apicultural Research 56:354365

112. Gerard, F., D. Hooftman, F. van Langevelde, E. Veenendaal, S.M. White and J. Lloyd (2017) MODIS VCF should not be used to detect discontinuities in tree cover due to binning bias – a comment on Hanan et al. (2014) and Staver and Hansen (2015). Global Ecology and Biogeography 26:854859

111. Essens, T., F. van Langevelde, R.A. Vos, C.A.M. van Swaay and M.F. Wallis de Vries (2017) Ecological determinants of butterfly vulnerability across the European
continent. Journal of Insect Conservation 21:439–450

110. Van Langevelde, F., R.H.A. van Grunsven, E.M. Veenendaal and T.P.M. Fijen (2017) Artificial night lighting inhibits feeding in moths. Biology Letters 13:20160874

109. Sianga, K., M. van Telgen, J. Vrooman, R. Fynn and F. van Langevelde (2017) Spatial refuges buffer landscapes against homogenization and degradation by large herbivore populations and facilitate vegetation heterogeneity. Koedoe 59:a1434

108. Huang, Z.X.Y., F. van Langevelde, K.J. Honer, M. Naguib and W.F. de Boer (2017) Regional level risk factors associated with the occurrence of African swine fever in West and East Africa. Parasites & Vectors 10:16

107. Huang, Z.Y.X, Y. Yu, F. van Langevelde and W.F. de Boer (2017) Does the dilution effect generally occur in animal diseases? Parasitology 144:823–826

106. Wynhoff, I., A.M. Kolvoort, C.F. Bassignana, M.P. Berg and F. van Langevelde (2017) Fen meadows on the move for the conservation of Maculinea (Phengaris) teleius butterflies. Journal of Insect Conservation 21:379–392

105. Nkoba, K., S.K. Raina and F. van Langevelde (2017) Impact of habitat degradation on species diversity and nest abundance of five African stingless bee species in a tropical rainforest of Kenya. International Journal of Tropical Insect Sciences 37:189–197


104. Wato, Y.A., I.M.A. Heitkönig, S.E. van Wieren, G. Wahungu, H.H.T. Prins and F. van Langevelde (2016) Prolonged drought results in starvation of African elephant (Loxodonta africana). Biological Conservation 203:89–96

103. Nkoba, K., S.K. Raina and F. van Langevelde (2016) A vertical compartmented hive design effective to reduce post-harvest colony loss in Afrotropical stingless bee species (Apidae: Meliponinae). International Journal of Development Research 6: 9026-9034

102. Van Langevelde, F., Z.K. Tessema, W.F. de Boer and H.H.T. Prins (2016) Soil seed bank dynamics under the influence of grazing as alternative explanation for herbaceous vegetation transitions in semi-arid rangelands. Ecological Modelling 337:253–261

101. Kölzsch, A., M. Neefjes, J. Barkway, G.J.D.M. Muskens, F. van Langevelde, W.F. de Boer, H.H.T. Prins, B.H. Cresswell and B.A. Nolet (2016) Neckband or backpack? Differences in tag design and their effects on GPS/accelerometer tracking results in large waterbirds. Animal Biotelemetry 4:13

100. Van der Waal, C., H. de Kroon, F. van Langevelde, W.F. de Boer, I.M.A. Heitkönig, R. Slotow, Y. Pretorius and H.H.T. Prins (2016) Scale-dependent bi-trophic interactions in a semi-arid savanna: how herbivores eliminate benefits of nutrient patchiness to plants. Oecologia 181:1173–1185

99. Cardoso, A.W., J.A. Medina-Vega, Y. Malhi, S. Adu-Bredu, G.K.D. Ametsitsi, G. Djagbletey, F. van Langevelde, E. Veenendaal and I. Oliveras (2016) Winners and losers: tropical forest tree seedling survival across a West African forest-savanna transition. Ecology and Evolution 6:3417–3429

98. Huang, Z.Y.X., F. van Langevelde, A. Estrada-Peña, G. Suzán and W.F. de Boer (2016) The diversity-disease relationship: evidence for and criticisms of the dilution effect. Parasitology 143:1075–1086

97. Weterings, M.J.A., M. Zaccaroni, N. van der Koore, L.M. Zijlstra, H. Kuipers, F. van Langevelde and S.E. van Wieren (2016) Strong reactive movement response of the medium-sized European hare to elevated predation risk in short vegetation. Animal Behaviour 115:107–114

96. Mwakiwa, E., J.W. Hearne, J.D. Stigter, W.F. de Boer, M. Henley, R. Slotow, F. van Langevelde, M. Peel, C.C. Grant and H.H.T. Prins (2016) Optimization of net returns from wildlife consumptive and non-consumptive uses by game reserve management. Environmental Conservation 43:128–139

95. Tomlinson, K.W., F. van Langevelde, D. Ward, H.H.T. Prins, S. de Bie, B. Vosman, E.V.S.B. Sampaio and F.J. Sterck (2016) Defence against vertebrate herbivores trades off into architectural and low nutrient strategies amongst savanna Fabaceae species. Oikos 125:126–136


94. Blanken, L.J., F. van Langevelde and C. van Dooremalen (2015) Interaction between Varroa destructor and imidacloprid reduces flight capacity of honeybees. Proceedings of the Royal Society B 282:20151738

93. Van Langevelde, F. (2015) Modelling the negative effects of landscape fragmentation on habitat selection. Ecological Informatics 30:271–276

92. De Weerd, N., F. van Langevelde, H. van Oeveren, B.A. Nolet, A. Kölzsch, H.H.T Prins and W.F. de Boer (2015) Deriving animal behaviour from high-frequency GPS: tracking cows in open and forested habitat. Plos ONE 10:e0129030

91. Ribeiro, L.C., E.R.M. Barbosa, F. van Langevelde and F. Borghetti (2015) The importance of seed biomass for the tolerance to heat shock of savanna and forest tree species. Journal of Vegetation Science 26:1102–1111

90. Meyer, N., H.J. Esser, R. Moreno, F. van Langevelde, Y. Liefting, D. Ros Oller, C.B.F. Vogels, A.D. Carver, C.K. Nielsen and P.A. Jansen (2015) An assessment of the terrestrial mammal communities in forests of Central Panama, using camera-trap surveys. Journal for Nature Conservation 26:28–35

89. Huang, Z.Y.X., F. van Langevelde, H.H.T. Prins and W.F. de Boer (2015) Dilution versus facilitation: impact of connectivity on disease risk in metapopulations. Journal of Theoretical Biology 376:66–73

88. Wynhoff, I., R.B. Bakker, B. Oteman, P.S. Arnaldo and F. van Langevelde (2015) Phengaris (Maculinea) alcon butterflies deposit their eggs on tall plants with many large buds in the vicinity of Myrmica ants. Insect Conservation and Diversity 8:177–188

87. Van de Poel, J.L., J. Dekker and F. van Langevelde (2015) Dutch hedgehogs (Erinaceus europaeus) are nowadays mainly found in urban areas, possibly due to the negative effects of badgers (Meles meles). Wildlife Biology 21:51–55

86. Hilbers, J.P., F. van Langevelde, H.H.T. Prins, C.C. Grant, M. Peel, M.B. Coughenour, H.J. de Knegt, R. Slotow, I.P.J. Smit, G.A. Kiker and W.F. de Boer (2015) Modeling elephant-mediated cascading effects of water point closure. Ecological Applications 25:402–415


85. Arnaldo, P.S., D. Gonzalez, I. Oliveira, F. van Langevelde and I. Wynhoff (2014) Influence of host plant phenology and oviposition date on the oviposition pattern and offspring performance of the butterfly Phengaris alcon. Journal of Insect Conservation 18:1115–1122

84. Thakur, M.P., M.P. Berg, N. Eisenhauer and F. van Langevelde (2014) Disturbance–diversity relationships for soil fauna are explained by faunal community biomass in a salt marsh. Soil Biology & Biochemistry 78:30–37

83. Tomlinson, K.W., L. Poorter, F. Bongers, F. Borghetti, L. Jacobs and F. van Langevelde (2014) Relative growth rate variation of evergreen and deciduous savanna tree species is driven by different traits. Annals of Botany 114:315–324

82. Barbosa, E.R.M., K.W. Tomlinson, L.G. Carvalheiro, K. Kirkman, S. de Bie, H.H.T. Prins and F. van Langevelde (2014) Short-term effect of nutrient availability and rainfall distribution on biomass production and leaf nutrient content of savanna tree species. Plos ONE 9:e92619

81. Comor, V., M.P. Thakur, M.P. Berg, S. de Bie, H.H.T. Prins and F. van Langevelde (2014) Productivity affects the density – body mass relationship of soil fauna communities. Soil Biology & Biochemistry 72:203–211

80. Van Langevelde, F., C. de Groot, T.A. Groen, I.M.A. Heitkönig and I. Gaigher (2014) Effect of patches of woody vegetation on the role of fire in tropical grasslands and savannas. International Journal of Wildland Fire 23:410–416

79. Huang, Z.Y.X., C. Xua, F. van Langevelde, H.H.T. Prins, K. Ben Jebara and W.F. de Boer (2014) Dilution effect and identity effect by wildlife in the persistence and recurrence of bovine tuberculosis. Parasitology 141:981–987

78. Barbosa, E.R.M., F. van Langevelde, K.W. Tomlinson, L.M.G.R. Carvalheiro, K. Kirkman, S. de Bie and H.H.T. Prins (2014) Tree species from different functional groups respond differently to environmental changes during establishment. Oecologia 174:1345–1357

77. Shrestha A.K., S.E. van Wieren, F. van Langevelde, A. Fuller, R.S. Hetem, L.C.R. Meyer, S. de Bie and H.H.T. Prins (2014) Larger antelopes are sensitive to heat stress throughout all seasons but smaller antelope only during summer in an African semi-arid environment. International Journal of Biometeorology 58:41–49


76. Jaarsma, C.F., F. van Langevelde and R. Beunen (2013) Landscape Ecology and Rural Roads: Traffic Calming for improving both landscape and wildlife? Ekológia 32:352–360

75. Huang, Z.Y.X., W.F. de Boer, F. van Langevelde, C. Xu, K. Ben Jebara, F. Berlingieri and H.H.T. Prins (2013) Dilution effect in bovine tuberculosis: risk factors for regional disease occurrence in Africa. Proceedings of the Royal Society B 280:20130624

74. Tomlinson, K.W., F. van Langevelde, D. Ward, F. Bongers, D.A. da Silva, H.H.T. Prins, S. de Bie and F.J. Sterck (2013) Deciduous and evergreen trees differ in juvenile biomass allometries because of differences in allocation to root storage. Annals of Botany 112:575–587

73. Van Dooremalen, C., E. Stam, L. Gerritsen, B. Cornelissen, J.J.M. van der Steen, F. van Langevelde and T. Blacquière (2013) Interactive effect of reduced pollen availability and Varroa destructor infestation limits growth and protein content of young honey bees. Journal of Insect Physiology 59:487–493

72. Tomlinson, K.W., L. Poorter, F. Sterck, F. Borghetti, D. Ward, S. de Bie and F. van Langevelde (2013) Leaf adaptations of evergreen and deciduous trees of semi-arid and humid savannas on three continents. Journal of Ecology 101:430–440

71. De Boer, W.F., F. van Langevelde, H.H.T. Prins, P.C. de Ruiter, J. Blanc, M.J.P. Vis, K.J. Gaston and I. Douglas Hamilton (2013) Understanding spatial differences in African elephant densities and occurrence, a continent-wide analysis. Biological Conservation 159:468–476

70. Huang, Z.Y.X., W.F. de Boer, F. van Langevelde, V. Olson, T.M. Blackburn and H.H.T. Prins (2013) Species’ life-history traits explain interspecific variation in reservoir competence: a possible mechanism underlying the dilution effect. PLoS ONE 8:e54341

69. Mwakiwa, E., W.F. de Boer, J.W. Hearne, R. Slotow, F. van Langevelde, M. Peel, C.C. Grant, Y. Pretorius, J.D. Stigter, A.K. Skidmore, I.M.A. Heitkönig, H.J. de Knegt, E.M. Kohi, N. Knox and H.H.T. Prins (2013) Optimization of wildlife management in a large game reserve through waterpoints manipulation: a bio-economic analysis. Journal of Environmental Management 114:352–361


68. Pretorius, Y., J.D. Stigter, W.F. de Boer, S.E. van Wieren, C.B. de Jong, H.J. de Knegt, C.C. Grant, I. Heitkönig, N. Knox, E. Kohi, E. Mwakiwa, M.J.S. Peel, A.K. Skidmore, R. Slotow, C. van der Waal, F. van Langevelde and H.H.T. Prins (2012) Diet selection of African elephant over time shows changing optimization currency. Oikos 121:2110–2120

67. Tomlinson, K.W., F.J. Sterck, F. Bongers, D.A. da Silva, E.R.M. Barbosa, D. Ward, F.T. Bakker, M. van Kaauwen, H.H.T. Prins, S. de Bie and F. van Langevelde (2012) Biomass partitioning and root morphology of savanna trees across a water gradient. Journal of Ecology 100:1113–1121

66. Jansen, S.H.D.R., M. Holmgren, F. van Langevelde and I. Wynhoff (2012) Resource use of specialist butterflies in agricultural landscapes: conservation lessons from the butterfly Phengaris (Maculinea) nausithous. Journal of Insect Conservation 16:921–930

65. Ahrestani, F., F. van Langevelde, I.M.A. Heitkönig and H.H.T. Prins (2012) Contrasting timing of parturition of chital (Axis axis) and gaur (Bos gaurus) in tropical South India – The role of body mass and seasonal forage quality. Oikos 121:1300–1310

64. Van Dooremalen, C., L. Gerritsen, B. Cornelissen, J.J.M. van der Steen, F. van Langevelde and T. Blacquière (2012) Winter survival of individual honey bees and honey bee colonies depends on level of Varroa destructor infestation. PLoS ONE 7:e36285

63. Kos, M., A.J. Hoetmer, Y. Pretorius, W.F. de Boer, H. de Knegt, C.C. Grant, E. Kohi, B. Page, M. Peel, R. Slotow, C. van der Waal, S.E. van Wieren, H.H.T. Prins and F. van Langevelde (2012) Seasonal diet change in elephant and impala in a mopane woodland. European Journal of Wildlife Research 58:279–287

62. Shrestha A.K., S.E. van Wieren, F. van Langevelde, A. Fuller, R.S. Hetem, L.C.R. Meyer, S. de Bie and H.H.T. Prins (2012) Body temperature variation of South African antelopes in two climatically contrasting environments. Journal of Thermal Biology 37:171–178


61. Ahrestani, F.S., I.M.A. Heitkonig, F. van Langevelde, V. Srinivas, M.D. Madhusudan and H.H.T. Prins (2011) Moisture and nutrients determine the distribution and richness of India’s large herbivore species assemblage. Basic and Applied Ecology 12:634–642

60. Van Langevelde, F., W.F. de Boer, H.J. de Knegt and H.H.T. Prins (2011) Emergent properties of the Lévy walk are not adaptive strategies. Science E-letter

59. Van Langevelde, F., J.A. Ettema, M. Donners, M.F. WallisDeVries and D. Groenendijk (2011) Effect of spectral composition of artificial light on the attraction of moths. Biological Conservation 144:2274–2281

58. Van Langevelde, F., K. Tomlinson, E.R.M. Barbosa, S. de Bie, H.H.T. Prins and SI. Higgins (2011) Understanding tree-grass co-existence and impacts of disturbance and resource variability in savannas. In: M.J. Hill and N.P. Hanan (eds.), Ecosystem function in savannas. Measurement and modeling at landscape to global scales. CRC Press, Boca Raton, FL, USA, pp. 257–271

57. Jaarsma, C.F. and F. van Langevelde (2011) Effects of scale and efficiency of rural traffic calming on safety, accessibility and wildlife. Transportation Research D 16:486–491

56. Groen, T.A., F. van Langevelde, C.A.D.M. van de Vijver, A.L. de Raad, J. de Leeuw and H.H.T. Prins (2011) A continental analysis of correlations between tree patterns in African savannas and human and environmental variables. Journal of Arid Environments 75:724–733

55. Van der Waal, C., H. de Kroon, I.M.A. Heitkönig, W.F. de Boer, A.K. Skidmore, F. van Langevelde, R. Slotow, R.C. Grant, M. Peel, E.M. Kohi, H.J. de Knegt and H.H.T. Prins (2011) Scale of nutrient patches mediates resource partitioning between trees and grasses in a semi-arid savanna. Journal of Ecology 5:1124–1133

54. Van Langevelde, F. and C.J. Grashof-Bokdam (2011) Modelling the effect of intersections in linear habitat on spatial distribution and local population density. International Journal of Geographical Information Science 25:367–378

53. Wynhoff, I., R. van Gestel, C. van Swaay and F. van Langevelde (2011) Not only the butterflies: Managing ants on road verges to benefit Phengaris (Maculinea) butterflies. Journal of Insect Conservation 15:189–206

52. De Knegt, H.J., F. van Langevelde, A.K. Skidmore, A. Delsink, R. Slotow, S. Henley, G. Bucini, W.F. de Boer, M. Coughenour, C.C. Grant, I.M.A. Heitkönig, M. Henley, N.M. Knox, E.M. Kohi, E. Mwakiwa, B.R. Page, M. Peel, Y. Pretorius, S.E. van Wieren and H.H.T. Prins (2011) The spatial scaling of habitat selection by African elephants. Journal of Animal Ecology 80:270–281

51. Heuermann, N., F. van Langevelde, S.E. van Wieren and H.H.T. Prins (2011) Increased searching and handling effort in tall swards lead to a Type IV functional response in small grazing herbivores. Oecologia 166:659–669

50. Van der Waal, C., A. Kool, S.S. Meijer, H. de Kroon, E. Kohi, I.M.A. Heitkönig, W.F. de Boer, F. van Langevelde, C.C. Grant, M.J.S. Peel, R. Slotow, H.J. de Knegt and H.H.T. Prins (2011) Large herbivores may alter vegetation structure of semi-arid savannas through soil nutrient mediation. Oecologia 165:1095–1107


49. De Boer, W.F., M.J.P. Vis, H.J. De Knegt, C. Rowles, E.M. Kohi, F. van Langevelde, M. Peel, Y. Pretorius, A.K. Skidmore, R. Slotow, S.E. van Wieren and H.H.T. Prins (2010) Spatial distribution of lion kills determined by the water dependency of prey species. Journal of Mammalogy 91:1280–1286

48. Jonker, R.M., G. Eichhorn, F. van Langevelde and S. Bauer (2010) Predation danger can explain changes in timing of migration: The case of the Barnacle Goose. PLoS ONE 5:e11369

47. De Knegt, H J., F. van Langevelde, M. B. Coughenour, A.K. Skidmore, W.F. de Boer, I.M.A. Heitkönig, N.M. Knox, R. Slotow, C. van der Waal and H.H.T. Prins (2010) Spatial autocorrelation and the scaling of species-environment relationships. Ecology 91:2455–2465


46. Okita-Ouma, B., R. Amin, F. van Langevelde and N. Leader-Williams (2009) Density dependence and population dynamics of black rhinos (Diceros bicornis michaeli) in Kenya’s rhino sanctuaries. African Journal of Ecology 48:791–799

45. Voeten, M.M., C.A.D.M. van de Vijver, H. Olff and F. van Langevelde (2009) Possible causes of decreasing migratory ungulate populations in an East African savanna after restrictions in their seasonal movements. African Journal of Ecology 48:169–179

44. Kohi, E.M., W.F. de Boer, M. Slot, S.E. van Wieren, J.G. Ferwerda, C.C. Grant, I.M.A. Heitkönig, N. Knox, F. van Langevelde, M. Peel, R. Slotow, C. van der Waal and H.H.T. Prins (2009) Effects of simulated browsing on growth and leaf chemical properties in Colophospermum mopane saplings. African Journal of Ecology 48:190–196

43. Van Langevelde, F. and C.F. Jaarsma (2009) Modelling the effect of traffic calming on local animal population persistence. Ecology and Society 14:39

42. Van Langevelde, F. and I. Wynhoff (2009) What limits the spread of two congeneric butterfly species after their reintroduction: quality or spatial arrangement of habitat? Animal Conservation 12:540–548

41. Huisman, T.J., F. van Langevelde and W.F. de Boer (2009) Local positive feedback and the persistence and recovery of fringe Avicennia marina (Forssk.) vierh. mangroves. Wetlands Ecology and Management 17:601–611

40. Van der Waal, C., H. de Kroon, W.F. de Boer, I.M.A. Heitkonig, A.K. Skidmore, H.J. de Knegt, F. van Langevelde, S.E van Wieren, C.C. Grant, B.R. Page, R. Slotow, E.M. Kohi, E. Mwakiwa and H.H.T. Prins (2009) Water and nutrient availability influence herbaceous competition effects on establishing woody plants in a semi arid savanna. Journal of Ecology 97:430–439

39. Hengeveld, G.M., F. van Langevelde, T.A. Groen and H.J. de Knegt (2009) Optimal foraging for multiple resources in several food species. American Naturalist 174:102–110

38. Van Langevelde, F., C. van Dooremalen and C.F. Jaarsma (2009) Traffic mortality and the role of minor roads. Journal of Environmental Management 90:660–667


37. Wynhoff, I., M. Grutters and F. van Langevelde (2008) Looking for the ants: selection of oviposition sites by two myrmecophilous butterfly species. Animal Biology 58:371–388

36. Groen, T.A., F. van Langevelde, C.A.D.M. van de Vijver, N. Govender and H.H.T. Prins (2008) Soil clay content and fire frequency affect clustering in savanna trees in South African savannas. Journal of Tropical Ecology 24:269–279

35. De Knegt, H., T.A. Groen, C.A.D.M. van de Vijver, H.H.T. Prins and F. van Langevelde (2008) Herbivores as architects of savannas: inducing and modifying spatial vegetation patterning. Oikos 117:543–554

34. Van Langevelde, F., M. Drescher, I.M.A. Heitkönig and H.H.T. Prins (2008) Modelling instantaneous intake rate of herbivores as function of forage quality and mass: effects on facilitative and competitive interactions between differently sized herbivores. Ecological Modelling 213:273–284

33. Prins, H.H.T. and F. van Langevelde (2008) Prospects for further developments of resource ecology. In: H.H.T. Prins and F. van Langevelde (eds.), Resource Ecology. Spatial and temporal aspects of foraging. Springer, Dordrecht, The Netherlands, pp. 267–271

32. Prins, H.H.T. and F. van Langevelde (2008) Assembling a diet from different places. In: H.H.T. Prins and F. van Langevelde (eds.), Resource Ecology. Spatial and temporal dynamics of foraging. Springer, Dordrecht, The Netherlands, pp. 129–155

31. Prins, H.H.T., A.M.H. Brunsting and F. van Langevelde (2008) Comments on “The relevance of key resource areas for large-scale movements of livestock”. In: H.H.T. Prins and F. van Langevelde (eds.), Resource Ecology. Spatial and temporal aspects of foraging. Springer, Dordrecht, The Netherlands, pp. 233–236

30. Van Langevelde, F. and Prins, H.H.T. (2008) Introduction on resource ecology. In: H.H.T. Prins and F. van Langevelde (eds.), Resource Ecology. Spatial and temporal aspects of foraging. Springer, Dordrecht, The Netherlands, pp. 1–6

29. Prins, H.H.T., W.F. de Boer and F. van Langevelde (2008) Comments on “Resource distribution and dynamics: mapping herbivore resources”. In: H.H.T. Prins and F. van Langevelde (eds.), Resource Ecology. Spatial and temporal aspects of foraging. Springer, Dordrecht, The Netherlands, pp. 79–80

28. Van Wieren, S.E. and F. van Langevelde (2008) Structuring of herbivore communities: the role of habitat and diet. In: H.H.T. Prins and F. van Langevelde (eds.), Resource Ecology. Spatial and temporal dynamics of foraging. Springer, Dordrecht, The Netherlands, pp. 237–262


27. De Knegt, H.J., G.M. Hengeveld, F. van Langevelde, W.F. de Boer and K.P. Kirkman (2007) Patch density determines movement patterns and foraging efficiency of large herbivores. Behavioural Ecology 18:1065–1072

26. Mourik, A.A., F. van Langevelde, E. van Tellingen, I.M.A. Heitkönig and I. Gaigher (2007) Stability of wooded patches in a South African nutrient-poor grassland: do nutrients, fire or herbivores limit their expansion? Journal of Tropical Ecology 23:529–537

25. Van Langevelde, F. and H.H.T. Prins (2007) Resilience and restoration of soft-bottom near-shore ecosystems. Hydrobiologia 591:1–4

24. Jaarsma, C.F., F. van Langevelde, J.M. Baveco, M. van Eupen and J. Arisz (2007) Model for rural transportation planning considering simulating mobility and traffic kills in the badger Meles meles. Ecological Informatics 2:73–82

23. Jansen, A.W.A.M., F. van Langevelde, W. F. de Boer and K.P. Kirkman (2007) Optimisation or satiation, testing diet selection rules in goats. Small Ruminant Research 73:160–168

22. Van Langevelde, F., C. van Dooremalen and C.F. Jaarsma (2007) Traffic mortality, analysis and mitigation. Effects of road, traffic, vehicle and species characteristics. In: S.K. Hong, N. Nakagoshi, B. Fu and Y. Morimoto (eds.), Landscape ecological applications in man-influenced areas – linking man and nature systems. Springer, Dordrecht, The Netherlands, pp. 253–272


21. Jaarsma, C.F., F. van Langevelde and H. Botma (2006) Flattened fauna and mitigation: traffic victims related to road, traffic, vehicle, and species characteristics. Transportation Research Part D 11:264–276

20. Kleijn, D. and F. van Langevelde (2006) Interacting effects of landscape context and habitat quality on flower visiting insects in agricultural landscapes. Basic and Applied Ecology 7:201–214


19. Van ’t Hof, P. and F. van Langevelde (2005) Reintroduction of the otter in the Netherlands: implementation of international guidelines in pre-release phase. Lutra 48:135–137

18. Van Gemerden, B.S, H. Olff, R.S. Etienne, P.W.F.M. Hommel and F. van Langevelde (2005) Reconciling methodologically different biodiversity assessments of species-rich tropical rain forests. Ecological Applications 15:1747–1760

17. Grashof-Bokdam, C.J. and F. van Langevelde (2005) Green veining: landscape determinants of biodiversity in European agricultural landscapes. Landscape Ecology 20:417–439


16. Van ’t Hof, P. and F. van Langevelde (2004) Reintroduction of otter (Lutra lutra) in The Netherlands meets international guidelines. Lutra 47:127–132

15. Van Langevelde, F. and C.F. Jaarsma (2004) Using traffic flow theory to model traffic mortality in mammals. Landscape Ecology 19:895–907

14. Stigter, J.D. and F. van Langevelde (2004) Optimal harvesting in a two-species model under critical depensation. The case of optimal harvesting in semi-arid grazing systems. Ecological Modelling 179:153–161


13. Van Langevelde, F., C.A.D.M. van de Vijver, L. Kumar, J. van de Koppel, N. de Ridder, J. van Andel, A.K. Skidmore, J. Hearne, L. Stroosnijder, W.J. Bond, H.H.T. Prins and M. Rietkerk (2003) Effects of fire and herbivory on the stability of savanna ecosystems. Ecology 84:337–350

12. Drescher, M. and F. van Langevelde (2003) Dynamics in herbivore aggregation resulting from seasonal changes in forage resource quality and production – a modelling approach. African Journal of Range & Forage Science 20:72


11. Rietkerk, M., M. Boerlijst, F. van Langevelde, R. HilleRisLambers, J. van de Koppel, L. Kumar, H.H.T. Prins and A. de Roos (2002) Self-organization of vegetation in arid ecosystems. American Naturalist 160:524–530

10. Van Langevelde, F., G.D.H. Claassen and A.G.M. Schotman (2002) Two strategies for conservation planning in human-dominated landscapes. Landscape and Urban Planning 58:281–296

9. Rietkerk, M., T. Ouedraogo, L. Kumar, S. Sanou, F. van Langevelde, A. Kiema, J. van de Koppel, J. van Andel, J. Hearne, A.K. Skidmore, N. de Ridder, L. Stroosnijder and H.H.T. Prins (2002) Fine-scale spatial distribution of plants and resources on a sandy soil in the Sahel. Plant and Soil 239:69–77

8. Kumar, L., M. Rietkerk, F. van Langevelde, J. van de Koppel, J. van Andel, J. Hearne, N. de Ridder, L. Stroosnijder, A.K. Skidmore and H.H.T. Prins (2002) Relationship between vegetation growth rates at the onset of the wet season and soil type in the Sahel of Burkina Faso: implications for resource utilisation at large scales. Ecological Modelling 149:143–152

7. Rietkerk, M., J. van de Koppel, L. Kumar, F. van Langevelde and H.H.T. Prins (2002) The ecology of scale. Ecological Modelling 149:1–4

6. Van de Koppel, J., M. Rietkerk, F. van Langevelde, L. Kumar, C.A. Klausmeier, J.M. Fryxell, J. Hearne, J. van Andel, N. de Ridder, A.K. Skidmore, L. Stroosnijder and H.H.T. Prins (2002) Spatial heterogeneity and irreversible vegetation change in semi-arid grazing systems. American Naturalist 159:209–218

before 2001

5. Van Langevelde, F. (2000) Scale of habitat connectivity and colonization in fragmented nuthatch populations. Ecography 23:614–622

4. Van Langevelde, F., A.G.M. Schotman, G.D.H. Claassen and G.A. Sparenburg (2000) Competing land uses in the reserve site selection problem. Landscape Ecology 15:243–256

3. Van Langevelde, F., W.G.M. van der Knaap and G.D.H. Claassen (1998) Comparing connectivity in landscape networks. Environment and Planning B 25:849–863

2. Van Langevelde, F. and C.F. Jaarsma (1997) Habitat fragmentation, the role of minor rural roads and their traversability. In: C. Kanters, A. Piepers and D. Hendriks-Heersma (eds.), Habitat Fragmentation & Infrastructure, pp. 171–182

1. Van Langevelde, F. (1994) Conceptual integration of landscape planning and landscape ecology, with a focus on the Netherlands. In: E.A. Cook and H.N. van Lier (eds.), Landscape Planning and Ecological Networks. ISOMUL 6F, Elsevier, Amsterdam, pp. 27–69

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